G. variabilis Davis and Pellmyr
Adult Characteristics
The most variable of all Greya species. Wing expanse 12-18 mm. Body pale ochreous to dark brown. Forewings generally with a purplish brown base color, with many patches and streaks of pale ochreous in different patterns. In some individuals, light areas are so extensive as to anastomose. Rare individuals have completely tan colored forwings. Variation is distributed both within and among populations. Pollex of male valva sometimes the result of two or more fused spines.

Assorted G. variabilis from range of species.
Comparison with similar species
Different color forms are often very similar to G. pectinifera, but are readily told apart by the genitalia. In the male, pectinifera has a pectinifer rather than a pollex. In the female, variabilis has lost the signa whereas two prominent signa are present in pectinifera.No other species approaches the general habitus of G. variata. G. kononenkoi is said to differ in having perpendicularly oriented forewing patches, and by having the tergum and sternum of segment VIII of equal length (tergum>sternum in variabilis).
Host, oviposition, and larval feeding habits
Unknown, but individuals often kicked up from areas with Saxifraga spp. (Saxifragaceae).
Geographic Distribution
Widely distributed, from the Chukchi peninsula of northwestern Siberia, the Pribilof Islands, interior Alaska, and several Pleistocene refugia along the North American west coast, including the Queen Charlotte Islands, the Olympic peninsula, and portions of the coastal range of northern Oregon.
Habitat
In the northern portion of the range, the species occur in tundra habitats. In more southern localities, it occurs in moist coniferous forest at intermediate elevation.

Characteristic habitat for G. variabilis on Olympic peninsula, Washington, USA.
See also picture under G. punctiferella.
References
Davis, D.R., O. Pellmyr & J.N. Thompson. 1992. Biology and systematics
of Greya Busck and Tetragma n. gen. (Lepidoptera: Prodoxidae).
Smiths. Contrib. Zool. 524:1-88.
Kozlov, M.V. 1996. Incurvariidae and Prodoxidae (Lepidoptera)
from Siberia and the Russian Far East, with descriptions
of two new species. Ent. Fenn. 7:55-62.
Holotype in USNM.
About this page
Olle Pellmyr
E-mail: pellmyr@ctrvax.vanderbilt.edu.
Dept of Biology, Vanderbilt University, Box 1812-B
Nashville, TN 37235, USA
Page copyright © 1996 Olle Pellmyr
Title Illustration:
Adult from Pribilof Islands, Alaska, USA.
Tree of Life design and icons copyright © 1996 David Maddison and Wayne Maddison.